1- Assistant Professor, Research Center for Plant Sciences, Ferdowsi University of Mashhad, Mashhad, IRAN , jafarnabati@ferdowsi.um.ac.ir 2- Professor, Faculty of Agriculture and Research Center for Plant Sciences, Ferdowsi University of Mashhad, Mashhad, IRAN 3- Faculty of Agriculture, Ferdowsi University of Mashhad, Mashhad, IRAN